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This work was supported by the awards from the Biotechnology and Biological Sciences Research Council [grant numbers BB/G023972/1, BB/R013748/1, BB/L026848/1, BB/M018431/1, BB/PO16855/1, BB/M001806/1, BB/P010520/1]; the European Research Council FUTUREROOTS Advanced Investigator grant [grant number 294729]; Leverhulme Trust [grant number RPG-2016-409]; Royal Society [grant number WM130021, NA140281]; Newton International Fellowship (NF140287) and British Council Newton Bhabha (228144076). This work was also supported by funds from the University of Nottingham Future Food Beacon of Excellence Nottingham Research and PhD+ fellowship schemes; the Interuniversity Attraction Poles Program initiated by the Belgian Science Policy Office [P7/29]; the Swedish Governmental Agency for Innovation Systems (VINNOVA), and the Swedish Research Council (V.R.) to K.L. We also thank Roger Granbom (Swedish University of Agricultural Sciences) for technical assistance and Gabriel Castrillo for commenting on the manuscript text. Part of this work has been conducted at the Rice Functional Genomics REFUGE platform funded by Agropolis Fondation in Montpellier, France. We also thank DBT-CREST BT/HRD/03/01/2002. ; Peer reviewed ; Publisher PDF